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Who follows the rule of the bull" means "who acts like a bull," who is virile by nature, or, perhaps more precisely, who, as a rule, is fecundating.
In these hazy, lush works, it's as if mathematics itself has become a fecundating principle.
Less well known is the extent to which Buddhist and Christian cultures became mutually fecundating in Central Asia in the first centuries of the Common Era.