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Grudgers will groom anyone--with one critical exception: if a member does not reciprocate, unlike the Suckers, the offended Grudger will punish the Cheater by ostracizing him from the group.
What motivates the Grudgers to engage in the act of punishing offenders at a personal cost to themselves?
It is this motivation to take action against the Cheaters that ultimately distinguishes the Grudgers from the Suckers, and enables them (us) to co-exist in the large, mutually beneficial social networks we recognize as modern society.