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Runs were performed in which the allele was nearly recessive (h = 0.
The presence of background deleterious mutations slows the advance of an advantageous recessive allele (Table 1).
Although direct estimates of d are not available for other organisms, the ubiquity of inbreeding depression for many fitness components in a wide variety of animal and plant species (reviewed by Charlesworth and Charlesworth 1987) shows that deleterious partial recessives are common.
Because it is well known that many alleles affecting fitness components and other quantitative traits are partially recessive (Simmons and Crow 1977; Crow and Simmons 1983; Charlesworth and Charlesworth 1987), it is important to determine whether this large class of genes can explain the results of the experiments cited above.