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paradoxus seeds is apparently controlled by endogenous factors that can prevent germination such as hormone levels, afterripening requirements, or possibly light and temperature sensitivity (Nikolaeva 1977; Baskin & Baskin 1998).
It is most likely a combination of factors that led to the relatively high germination of achenes following warm pretreatment, but this period may have allowed or stimulated the process of afterripening.
The differences may be due to the additional afterripening of the Utah seeds, differences in preconditioning (Baskin and Baskin, 1973), or the two populations from different parts of the species range may be displaying adaptations to significant locally selective pressures (Thompson, 1981).