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The change of recessive allele frequency as the function of this frequency in the previous generation q and the selection coefficient s is calculated as (Crow & Kimura, 1970: eq.
The resulting selection coefficients (s = 1 - [omega]) indicate that immigrants invading mature demes face much stronger selection gradients than invaders of young deme s (Table 4).
This position is supported by the fact that estimates of the degree of inbreeding depression for fitness in Drosophila are in very good agreement with expectations based on the estimates of mutation rates, selection coefficients, and coefficients of dominance obtained from mutation-accumulation experiments (Simmons and Crow 1977; D.
In this way, selection coefficients and differentials may be directly compared between populations and among traits.
A second type of kin selection coefficient arises when a phenotype influences different components of fitness.
By chance, some genomes have more deleterious alleles and/or deleterious alleles with larger selection coefficients than other genomes, and the alleles of these genomes are more readily lost.
2] and the selection coefficient S, based on the number of a plant's progeny surviving to age 6 (Falconer 1981).
These arguments suggest that with respect to the decline in fitness due to mutation accumulation, obligately selfing populations with mutation rate [mu] and selection coefficient s should be roughly equivalent to clonal populations with mutation rate [mu]/2 and selection coefficient 2s.
where s is the selection coefficient against a homozygous mutation at a single locus, and h is the dominance coefficient such that the fitness of the heterozygote is reduced by hs (Kimura and Maruyama 1966).