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Since the best option for crop production, under stressful conditions, is to improve both yield and yield stability through indirect selection, variability and heritability of traits related to stress tolerance need to be studied.
Several scientists have proposed methods to predict direct and indirect selection responses (Falconer, 1989; Bourdon, 2000).
The broad sense heritability of medium to high magnitude for the majority of traits indicated that direct or indirect selection enables selection with satisfactory gains.
Therefore, selection for shell length or shell width can result in indirect selection for body weight or muscle weight.
This means that there is a form of indirect selection occurring with people buying standard annuities in the open market being healthier and living longer than may be have been expected in the past.