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Related to mirthfulness: insolence
See: jocular
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Some of them were accurate -- an "organ of mirthfulness", for example, towards the front of the head.
Jennifer Raimundo finds the key for her analysis of mirthfulness in Tolkien's "On Fairy-stories," linking laughter and humor to Escape, Recovery, and Consolation.
She teases him only from mirthfulness: "You think you are a sage, but I think you are a fool and it's all the fault of that guy called Master Kong [Confucius] for turning you into a blockhead" (Qu 1989, 389-90).