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Related to pollinates: Cross pollination
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Bees pollinate the fragrant, pink-and-white tubular blooms, which cluster atop waist-high plants.
More than 100 different species (kinds) of birds pollinate more than 1000 species of flowers on that continent.
Freund explained that symptoms increase around February of each year, when trees begin to pollinate, and don't get much better until June, when the last of the trees - the olive - pollinates.
The color, shape, and length of the spur makes the nectar deep inside accessible to certain animals, enabling columbines to specialize and consequently diversify based on what pollinates them.