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Pressure is then applied on the central cornea with the goniolens, which stretches the limbal scleral ring, causing the iris to flatten and the ciliary body and iris to rotate posteriorly, thus opening the angle.
By taking measurements of the bony circle called the "scleral ring" in their eye and comparing it to the eye socket size in 164 living lizards and birds as well as 33 dinosaur fossils, scientists could tell which ones saw in the dark.
(17,18) The avian sclera is composed of hyaline cartilage lined by dense connective tissue (18-20) and at the intersection of the conjunctiva-sclera is the ossicles scleral ring. (21)