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Divergence times were not evaluated below the subordinal level in McKenna and Farrell (2009) due to the lack of well-supported resolution at lower taxonomic levels in Polyphaga and beyond.
Studies focused primarily on reconstructing relationships in Holometabola (e.g., Whiting et al., 1997, 2002a,b; Wheeler et al., 2001) using 18S and/or 28S DNA sequence data give some insight into subordinal relationships in beetles.
Por tanto, un lexicon de patrones asociado a una FPS representara y organizara variantes y combinaciones tipicas de patrones en el nivel subordinal de categorizacion.